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Thread: North Italian genetic structure

  1. #1
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    North Italian genetic structure

    There is the common, widespread misconception that all North Italians are more "northern" than central Italians, and that they all have Germanic admixture from the Longobards. For instance, 23andme writes in their article on Italians:

    "In the early Middle Ages, Germanic invaders brought about the fall of the Western Roman Empire, and a northern European genetic signature persists in modern Italians to this day. This influence is strongest in the north, while southern Italians share a rich genetic heritage with island Greeks."

    Now check out this map showing the incidence of blond hair colour in Italy around 1860:

    800px-BiasuttiMappa.jpg

    The map is based on data gathered by Ridolfo Livi on army conscripts from 1859-1863. Hence it shows Italian structure at a time when it was perhaps at its peak; for sure it was more structured than it is at present, after the migration of many South Italians to the north, in search of jobs, and since then Italians have also become more mobile within the regions.

    We can see that the majority of the lowland parts of the Emilia-Romagna are as dark as the southern half of central Italy. There are two spots, one around Forli, the other around Mirandola, that are even as dark as Calabria, or Southern Apulia.

    In this version of the same map, the exact locations of the spots can be better discerned by help of the rivers:

    hrtsh.jpg

    But it would be foolish to think that these places are genetically similar to Central or South Italians.

    The little that has been published on autosomal DNA from the Emilia shows that at least the Emilia is clearly part of the North Italian cluster. From the Romagna I don't know any published autosomal samples. From what I gathered from PCAs from hobbyists and from analyses from DNA kits, at least part of the Romagna resembles central Italy. But probably not all of it, as my maternal grandfather's roots were from Cesena, and the signal that I inherited is North Italian-like.

    The thing is, the main structure of Italy, as I understand it, isn't the result of more or less northern/central European/IE steppe ancestry; instead, it's the result of more or less Bronze Age Anatolian ancestry, which peaks in the South and is weakest in the north.

    Understandably, I find the dark spot around Forli particularly interesting, because Cesena is just nearby.

    Benito Mussolini's mother was from Forli:

    https://it.wikipedia.org/wiki/File:R..._mussolini.jpg

    While his father was from nearby Predappio:

    https://it.wikipedia.org/wiki/File:A..._mussolini.jpg

    They both look pretty exotic from a central European perspective, and pretty "typically" Italian.

    As did the "duce" himself:

    https://it.wikipedia.org/wiki/File:B...gshot_1903.jpg

    https://it.wikipedia.org/wiki/File:B...ni_colored.jpg

    Also from Forli, the singer Alice:

    https://youtu.be/czA66jXjEVE

    Now don't tell me that these are Celts, or even Longobards, lol.

    So how can we explain this? At the dawn of history there were the Umbri and the Etruscans in the Romagna, followed by the Gaulish invasion. Taking into account how light haired the modern Tuscans and Umbrians are, and bearing in mind that, even if the Umbri and Etruscans didn't all flee from the Gauls, but mixed with them, the Gaulish admixture must have increased the incidence of light hair even further. The modern inhabitants of Forli therefore cannot be predominantly descended from these. I believe they are predominantly descended from settlers who arrived with the Romans. That is, ancient Italic central and South Italians. Because these still had little or no Bronze Age Anatolian / Greek admixture, modern people from Forli and the wider (lowland) Emilia-Romagna are nonetheless North Italian-like, not Central or Southern.

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  3. #2
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    The Germanic Admixture in most of North Italy is maximum 15%.

    If you check some of the north Italian samples on G25 and do a run with relevant ancient samples. You will soon find that they have majority Italic Admixture with low Anatolian/Levant and some Germanic/Possible Celtic.

    I got central Italian admixture and score very little if any ancient Greek/Levant/Anatolian.

    known Italian ancestry - 25-37.5% Tuscan;

    Target: Nino_scaled
    Distance: 2.8731% / 0.02873087
    60.4 Medieval-German
    28.0 Early-Latin
    11.6 Iron-Age-Saami


    Here are some North and Central Italian averages:

    Target: Italian_Lombardy
    Distance: 0.9070% / 0.00907037 | ADC: 0.25x RC
    58.0 Early-Latin
    28.2 Etruscan
    13.8 Imperial-Roman
    Maybe I would add a Slavic reference sample for the Veneto. Because 29% Germanic is unrealistic.

    Target: Italian_Veneto
    Distance: 0.9661% / 0.00966107 | ADC: 0.25x RC
    43.6 Early-Latin
    29.4 Medieval-German
    26.0 Imperial-Roman
    1.0 Etruscan
    Target: Italian_Tuscany
    Distance: 0.6935% / 0.00693546 | ADC: 0.25x RC
    42.6 Imperial-Roman
    23.8 Early-Latin
    18.0 Medieval-German
    15.6 Etruscan
    Target: Italian_Bergamo
    Distance: 1.0927% / 0.01092690 | ADC: 0.25x RC
    38.8 Etruscan
    32.8 Early-Latin

    15.6 Imperial-Roman
    12.8 Medieval-German

    The problem is that we lack North Italian antique samples. Both Cisapline Gauls and Iron age Italic tribes.

    Once we get those samples, The Germanic will decrease much.

    Code:
    Early-Latin:ITA_Rome_Latini_IA,0.127482,0.147252,0.033187,-0.016796,0.044008,-0.008646,-0.00376,-0.004846,0.026588,0.052666,-0.002761,0.015137,-0.036719,-0.008533,-0.009093,0.013392,0.016037,-0.004687,0.003897,0.004127,0.00262,-0.00272,0.001972,-0.007712,-0.008742
    Early-Latin:ITA_Ardea_Latini_IA,0.133173,0.156392,0.0445,-0.00969,0.044008,-0.004462,0.00846,0.003,0.024543,0.044101,-0.002598,0.012289,-0.022448,-0.009634,-0.005429,-0.005834,0.00352,0.003167,0.006034,-0.007379,-0.008859,0.005317,-0.000863,-0.006989,-0.001796
    Early-Latin:ITA_Prenestini_tribe_IA,0.12862,0.145221,0.04714,0.005491,0.046162,0.00251,-0.00282,0.002769,0.021475,0.039727,-0.002923,0.012289,-0.021407,-0.016239,0.012622,0.000398,-0.003129,0.003421,0.00176,-0.001251,0.004492,0.003586,-0.00456,-0.003976,0.000239
    Early-Latin:ITA_Proto-Villanovan,0.125205,0.138112,0.030547,-0.010982,0.023081,-0.011713,0.002115,-0.001846,0.006954,0.021139,0.008119,0.005845,-0.011596,-0.000826,-0.007465,-0.01432,-0.011604,0.005194,0.002765,-0.00988,-0.009733,0.001607,-0.002465,0.003976,-0.005508
    Early-Latin:ITA_Villanovan,0.12862,0.156392,0.042238,-0.024225,0.052933,-0.008367,-0.001645,-0.005307,0.023725,0.040274,0.006008,0.006894,-0.018285,-0.019405,-0.003529,0.008751,0.025686,0.002914,0.004651,0.004252,0.00574,0.006059,-0.006286,-0.010604,-0.002515
    
    Imperial-Roman:ITA_Rome_Imperial,0.1039821,0.1495156,-0.0235307,-0.0574065,0.0045265,-0.0204055,-0.0011946,-0.0051488,0.0006604,0.0196549,0.0034575,0.0025539,-0.0040602,-0.0014737,-0.0081715,-0.0014474,0.0035992,0.000454,0.0012178,-0.0032854,-0.0025579,0.0020454,-0.0006985,-0.0004845,0.0004141
    Etruscan:ITA_Etruscan,0.126913,0.153853,0.034695,-0.014212,0.0417,-0.005299,0.0024675,-0.002769,0.019839,0.0343515,-0.0023545,0.014237,-0.014792,-0.004335,-0.00475,-0.007823,-0.0028035,0.0052575,0.011564,-0.001188,-0.002558,0.003771,-6.15e-05,-0.0004215,-0.002694
    Medieval-German:DEU_MA,0.1223596,0.1303939,0.061169,0.048773,0.039792,0.0199408,0.010975,0.0052151,0.0013295,-0.0024966,-0.003735,0.001109,-0.0091576,-0.0038398,0.0161643,-0.0008352,-0.0133511,0.0032684,0.0041354,0.0040271,0.0060019,0.0037342,-0.0007273,0.011146,-0.0004429
    Hidden Content


    Target: Nino_scaled
    Distance: 1.6700% / 0.01669968 | R3P
    53.6 Goth_Kowalewko = Germanic
    39.2 Levant_LBN_MA_o6 = South Euro
    7.2 RUS_Khaptsagai_IA = Steppe + Siberian

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    Quote Originally Posted by Nino90 View Post







    Code:
    Early-Latin:ITA_Rome_Latini_IA,0.127482,0.147252,0.033187,-0.016796,0.044008,-0.008646,-0.00376,-0.004846,0.026588,0.052666,-0.002761,0.015137,-0.036719,-0.008533,-0.009093,0.013392,0.016037,-0.004687,0.003897,0.004127,0.00262,-0.00272,0.001972,-0.007712,-0.008742
    Early-Latin:ITA_Ardea_Latini_IA,0.133173,0.156392,0.0445,-0.00969,0.044008,-0.004462,0.00846,0.003,0.024543,0.044101,-0.002598,0.012289,-0.022448,-0.009634,-0.005429,-0.005834,0.00352,0.003167,0.006034,-0.007379,-0.008859,0.005317,-0.000863,-0.006989,-0.001796
    Early-Latin:ITA_Prenestini_tribe_IA,0.12862,0.145221,0.04714,0.005491,0.046162,0.00251,-0.00282,0.002769,0.021475,0.039727,-0.002923,0.012289,-0.021407,-0.016239,0.012622,0.000398,-0.003129,0.003421,0.00176,-0.001251,0.004492,0.003586,-0.00456,-0.003976,0.000239
    Early-Latin:ITA_Proto-Villanovan,0.125205,0.138112,0.030547,-0.010982,0.023081,-0.011713,0.002115,-0.001846,0.006954,0.021139,0.008119,0.005845,-0.011596,-0.000826,-0.007465,-0.01432,-0.011604,0.005194,0.002765,-0.00988,-0.009733,0.001607,-0.002465,0.003976,-0.005508
    Early-Latin:ITA_Villanovan,0.12862,0.156392,0.042238,-0.024225,0.052933,-0.008367,-0.001645,-0.005307,0.023725,0.040274,0.006008,0.006894,-0.018285,-0.019405,-0.003529,0.008751,0.025686,0.002914,0.004651,0.004252,0.00574,0.006059,-0.006286,-0.010604,-0.002515
    
    Imperial-Roman:ITA_Rome_Imperial,0.1039821,0.1495156,-0.0235307,-0.0574065,0.0045265,-0.0204055,-0.0011946,-0.0051488,0.0006604,0.0196549,0.0034575,0.0025539,-0.0040602,-0.0014737,-0.0081715,-0.0014474,0.0035992,0.000454,0.0012178,-0.0032854,-0.0025579,0.0020454,-0.0006985,-0.0004845,0.0004141
    Etruscan:ITA_Etruscan,0.126913,0.153853,0.034695,-0.014212,0.0417,-0.005299,0.0024675,-0.002769,0.019839,0.0343515,-0.0023545,0.014237,-0.014792,-0.004335,-0.00475,-0.007823,-0.0028035,0.0052575,0.011564,-0.001188,-0.002558,0.003771,-6.15e-05,-0.0004215,-0.002694
    Medieval-German:DEU_MA,0.1223596,0.1303939,0.061169,0.048773,0.039792,0.0199408,0.010975,0.0052151,0.0013295,-0.0024966,-0.003735,0.001109,-0.0091576,-0.0038398,0.0161643,-0.0008352,-0.0133511,0.0032684,0.0041354,0.0040271,0.0060019,0.0037342,-0.0007273,0.011146,-0.0004429
    Neither of my parents, and therefore neither do I, have any Italian ancestors.

    Target: gixajo_scaled
    Distance: 2.1543% / 0.02154306

    84.2 Early-Latin
    9.0 Medieval-German
    6.8 Imperial-Roman

    Target: gixajo_dad_scaled
    Distance: 2.6721% / 0.02672113

    83.0 Early-Latin
    14.6 Imperial-Roman
    2.4 Etruscan

    Target: gixajo_mom_scaled
    Distance: 2.4229% / 0.02422862

    87.0 Early-Latin
    13.0 Medieval-German

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  7. #4
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    Quote Originally Posted by Simon_W View Post

    They both look pretty exotic from a central European perspective, and pretty "typically" Italian.
    I wouldn't get too much into that "phenotype" discussion pointing out northern Italians looking Italian vs central European bc it seems very subjective and unscientific. We didn't know what these people looked like 100% certain all we have is some snps to estimate relatively speaking. But it is true some provinces in the north are more typical genetically like their italian central parts... but the very northern parts bordering the Swiss do get an elevated Germanic admixture which isn't very surprising.


    Quote Originally Posted by mokordo View Post
    Neither of my parents, and therefore neither do I, have any Italian ancestors.

    Target: gixajo_scaled
    Distance: 2.1543% / 0.02154306

    84.2 Early-Latin
    9.0 Medieval-German
    6.8 Imperial-Roman

    Target: gixajo_dad_scaled
    Distance: 2.6721% / 0.02672113

    83.0 Early-Latin
    14.6 Imperial-Roman
    2.4 Etruscan

    Target: gixajo_mom_scaled
    Distance: 2.4229% / 0.02422862

    87.0 Early-Latin
    13.0 Medieval-German
    oh wow your distance is very close... it must be an overlapping issue bc those "early-latin" samples + etruscan are pretty similar to (eastern) iberian samples.

    compared to me:
    Target: Luso_scaled
    Distance: 4.1172% / 0.04117242
    78.0 Early-Latin
    13.0 Imperial-Roman
    9.0 Medieval-German

    -- these northern samples are very spanish-like:
    Villanova:

    Distance to: ITA_Villanovan
    0.03302067 Spanish_La_Rioja
    0.04462545 Spanish_Castello
    0.04551165 French_Corsica
    0.04682108 Spanish_Catalunya_Central
    0.04744870 Spanish_Baleares
    0.04778522 Spanish_Peri-Barcelona
    0.04804861 Spanish_Murcia



    this as well:

    Distance to: ITA_Prenestini_tribe_IA
    0.01431569 French_South
    0.01729684 Spanish_Aragon
    0.01775701 Spanish_Pirineu
    0.01786436 Spanish_Lleida
    0.01813429 Spanish_Castello


    Distance to: ITA_Ardea_Latini_IA
    0.03414474 Spanish_Terres_de_l'Ebre
    0.03481290 Spanish_Menorca
    0.03505798 Spanish_Pirineu
    0.03525011 Spanish_Baleares
    0.03555346 Spanish_Peri-Barcelona
    0.03584994 Spanish_Catalunya_Central
    0.03586453 Italian_Bergamo
    0.03666856 Italian_Trentino-Alto-Adige
    Last edited by Luso; 07-11-2021 at 12:03 AM.
    History of the Algarve: Hidden Content

    Paternal Lineage- R-Z225: São Bartolomeu de Messines- em Silves, Portugal

    Maternal Lineage: H1c3: Estrela de Monchique- em Silves, Portugal

    Silves, Portugal: Hidden Content

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    Ke5tEGd.jpg

    Here is a cool map I found a while ago(Cant remember where from)

    The names attached to the labels are from academic journals/reports which go into further detail about different roman migrations. Obviously as seen in the Roman Crossroads study there were genetic shifts after this period but it does give you a much clearer image of roman Italy(from an ethnic standpoint).

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    Quote Originally Posted by Rwilson98 View Post
    Here is a cool map I found a while ago(Cant remember where from)
    Indeed an interesting concept to look at the post Roman conquest shifts in ethnic composition trough veteran settlement. I would be very interested to find the original source which so far I was not able to locate despite the indication at the bottom right in addition to the title of the figure. I'm interested mostly in the Raetic, Venetic and Gallic areas. At least for Raetic I would expect some veteran settlement in the Athesis (Adige/Etsch) valley and even Isarcus (Isarco/Eisack) and Puster valley. But it all depends on the used sources (written/archeological) and the numbers in comparison.
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    Discussing North Italian genetic structure based on phenotype/lightness of features doesnt seem very helpful, you know.

    A model I've seen some people using for N. Italians is the Imperial_Roman + CHE_IA + Germanic_IA triad.
    I did a few changes to it but I'm not sure if it's good:



    Something wild is happening there with the Italian_Liguria sample I had to add the Guanche average only for the Ligurian one.
    Is the TUR_MLBA avg. a good proxy for bronze age anatolian input?
    If anyone has an average for Emilia-Romagna or Piedmont I'd be quite grateful if you could share them!
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    mtDNA 3x-GG H Reggio Calabria, Calabria, Italy
    Y-DNA 3x-GG R1b-U152 Modena, Emilia-Romagna, Italy
    Y-DNA 4x-GG E1b-V13 Reggio Calabria, Calabria, Italy
    Y-DNA 4x-GG R1b-U106 Venezia, Veneto, Italy

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    Quote Originally Posted by Simon_W View Post
    Now check out this map showing the incidence of blond hair colour in Italy around 1860:
    800px-BiasuttiMappa.jpg
    The map is based on data gathered by Ridolfo Livi on army conscripts from 1859-1863.
    I would be interested how Livi gathered and harmonized the data from the then different administrations unless French standardization was so dominant to facilitate this. See a map of the situation right before the conscripts time in 1843:


    Regarding the hair phenotype and with all the limited connection to the overall genetic admixture from the map I observe and would assume the North-Western part is strongly influenced by Celtic dominance (not sure how much blonde hair was present in the Alpine Gauls) as well as Germanic (Allemanic) incursions and later settlement (see Walser in Aosta XII-XIII century). In the historic Southern Tyrol area (now Trentino-Südtirol/Alto Adige) from what I see so far is strongly from Germanic settlements in the Middle Ages (from Bavaric, Frankish and Allemanic territory). The map lets suspect the same pattern also for the Cimbri/Zimbar area (language mainly derived from Medieval South Bavarian) near Vicenza (altopiano dei Sette Comuni / Asiago), near Verona (Tredici Comuni) and the Lusern village in Trentino (language still in use there). In the North-Eastern part it might be a mix of the Carinthian-Germanic influence as well as Slavic expansions with the latter probably stronger.
    Last edited by ChrisR; 07-11-2021 at 03:51 PM. Reason: correction
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    the recent paper with the bronze age samples also incl. a chart on hair and eye colors for a whole bunch of aDNA samples (incl. the RMPR) and contrary to popular believe traits like blond hair (also red) and blue eyes actually existed before the germanic invasions (imperial times) in samples fully comitted to the Med. and East-Med. clusters; and likewise blondism in late antiquity samples like san ercolano which are heavily central/south-italian like; so the whole looks department is not really such an indicator of north and south DNAwise

    the italian cline, as shown by the previous paper, is dependent on the gradient of italic_IA ancestry and future papers will show whether it is pre-roman italic or post-roman provincial (the gradient); north italy are *atleast two main clusters nw and ne and the closest clusters to central italians and south balkan (balkan1 = albanians)
    from raven et al (HDD 591,217 SNPs) CP/fineSTRUCTURE
     


    admixture wise there is no great diff. between north and central italians
    its mostly just isolation and drift (see Fst and IBD)

    *NITALY2: 2xTSI 28xEMI 20xLIG 15xLOM 29xPIE 1xTRE 1xVDA 5xVEN NITALY3: 1xLIG 22xLOM NITALY4: 2xLIG 5xTRE 11xVEN 14xFRI and the aostans actually branch with the italians (not french) guess that means their haplotype structure is more similar to the former than the latter (but acc. to Fst very drifted)
    GENO2.0 51SEURO 19WCEURO 13SCANDINAVIA 5ASIAMINOR 4EEURO 4GB/IRELAND 3ARABIA myOrigins 26ITA.PEN. 13GREECE&BALKANS 12SARDINIA 18GREATBRITAIN 14IRELAND 10C.EUROPE 8SCANDINAVIA DNA.Land 49NWEURO 27SEURO 13MED.ISLANDER 11SARDINIAN myHeritage 51.8NWEURO 33.2ITALIAN 7.9GREEK/S.ITALY 7.1BALKAN gencove 29NITALY 19EMED 15NBRITISLES 12SWEURO 10NCEURO 9SCANDINAVIA 6NEEURO GenePlaza 54.4NWEURO 37.6GRE/ALB 5.6WASIAN 2.4SWASIA LivingDNA 57.4S.GER 3.3NE.GER 25.8N.ITA 5S.ITA 4.3TUSCANY 2.5CYPRUS 1.7AEGEAN

  17. The Following 5 Users Say Thank You to alexfritz For This Useful Post:

     Alain (07-11-2021),  Cascio (08-26-2021),  Luso (07-11-2021),  Michalis Moriopoulos (07-11-2021),  peloponnesian (07-11-2021)

  18. #10
    Registered Users
    Posts
    366
    Sex
    Location
    Spain
    Ethnicity
    Español
    Nationality
    Española
    Y-DNA (P)
    E-V22 (E-FT156439)
    mtDNA (M)
    J1c5c1
    mtDNA (P)
    U6a3b1

    Spain Andalucia
    Distance to: hantro_scaled
    0.03426806 Early-Latin:ITA_Prenestini_tribe_IA
    0.04699128 Etruscan:ITA_Etruscan
    0.04828567 Early-Latin:ITA_Ardea_Latini_IA
    0.05797365 Early-Latin:ITA_Villanovan
    0.05898245 Early-Latin:ITA_Rome_Latini_IA
    0.05958917 Early-Latin:ITA_Proto-Villanovan
    0.08151249 Medieval-GermanEU_MA
    0.10879290 Imperial-Roman:ITA_Rome_Imperial

    Distance to: hantro_scaled
    0.03128993 88.20% Early-Latin:ITA_Prenestini_tribe_IA + 11.80% Imperial-Roman:ITA_Rome_Imperial
    0.03288391 83.80% Early-Latin:ITA_Prenestini_tribe_IA + 16.20% Early-Latin:ITA_Proto-Villanovan
    0.03365307 83.60% Early-Latin:ITA_Prenestini_tribe_IA + 16.40% Etruscan:ITA_Etruscan
    0.03396108 91.20% Early-Latin:ITA_Prenestini_tribe_IA + 8.80% Early-Latin:ITA_Villanovan
    0.03408722 6.80% Early-Latin:ITA_Rome_Latini_IA + 93.20% Early-Latin:ITA_Prenestini_tribe_IA
    0.03416057 96.40% Early-Latin:ITA_Prenestini_tribe_IA + 3.60% Medieval-GermanEU_MA
    0.03420319 5.80% Early-Latin:ITA_Ardea_Latini_IA + 94.20% Early-Latin:ITA_Prenestini_tribe_IA
    0.03886962 61.80% Early-Latin:ITA_Rome_Latini_IA + 38.20% Medieval-GermanEU_MA
    0.03946242 62.60% Early-Latin:ITA_Villanovan + 37.40% Medieval-GermanEU_MA
    0.03998535 72.40% Early-Latin:ITA_Ardea_Latini_IA + 27.60% Medieval-GermanEU_MA
    0.04078780 75.20% Etruscan:ITA_Etruscan + 24.80% Medieval-GermanEU_MA
    0.04579071 22.80% Early-Latin:ITA_Villanovan + 77.20% Etruscan:ITA_Etruscan
    0.04584330 40.60% Early-Latin:ITA_Ardea_Latini_IA + 59.40% Etruscan:ITA_Etruscan
    0.04588751 71.60% Early-Latin:ITA_Ardea_Latini_IA + 28.40% Early-Latin:ITA_Proto-Villanovan
    0.04599741 20.60% Early-Latin:ITA_Rome_Latini_IA + 79.40% Etruscan:ITA_Etruscan
    0.04622071 48.20% Early-Latin:ITA_Proto-Villanovan + 51.80% Early-Latin:ITA_Villanovan
    0.04689671 7.40% Early-Latin:ITA_Proto-Villanovan + 92.60% Etruscan:ITA_Etruscan
    0.04724907 90.80% Early-Latin:ITA_Ardea_Latini_IA + 9.20% Imperial-Roman:ITA_Rome_Imperial
    0.04735949 78.00% Early-Latin:ITA_Ardea_Latini_IA + 22.00% Early-Latin:ITA_Villanovan
    0.04742281 50.80% Early-Latin:ITA_Rome_Latini_IA + 49.20% Early-Latin:ITA_Proto-Villanovan
    0.04766163 18.20% Early-Latin:ITA_Rome_Latini_IA + 81.80% Early-Latin:ITA_Ardea_Latini_IA
    0.05284433 69.20% Early-Latin:ITA_Proto-Villanovan + 30.80% Medieval-GermanEU_MA
    0.05436255 39.20% Imperial-Roman:ITA_Rome_Imperial + 60.80% Medieval-GermanEU_MA
    0.05521152 46.00% Early-Latin:ITA_Rome_Latini_IA + 54.00% Early-Latin:ITA_Villanovan
    0.05671844 88.60% Early-Latin:ITA_Villanovan + 11.40% Imperial-Roman:ITA_Rome_Imperial
    Last edited by hantrolugharsts; 07-11-2021 at 01:12 PM.

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