Massive new paper from Allenthoft and Willerslev

[Trojet]

NEO806, Grotta Delle Mura, Monopoli, Italy_BronzeAge: J2b2a1a1a1a1a1b

Which corresponds to: J2b-L283>>Z638>Z1297>Z1295>Y21878

This should be a Iapygian (Paucetian?) sample, as it comes just south of Bari and the dating is listed as -1063 (BC).

Interesting to note, this seems to be the only sample from the region and timeframe in this paper, and turned out to be a J2b-L283

[Dewsloth]

No L238, DF19 or DF99 in the study's new samples, I think.
I note they have collected no samples from the Netherlands or northern Germany below the Jutland Peninsula, so where the cutoff is, I'm still not sure.

Geographically, the sampled skeletons cover a vast territory across Eurasia, from Lake Baikal to the
245 Atlantic coast, from Scandinavia to the Middle East, and they derive from a variety of contexts,
246 including burial mounds, caves, bogs and the seafloor (Supplementary Notes 6-7). Broadly, we can
247 divide our research area into three large regions: 1) central, western and northern Europe, 2) eastern
248 Europe including western Russia and Ukraine, and 3) the Urals and western Siberia. Our samples
249 cover many of the key Mesolithic and Neolithic cultures in Western Eurasia, such as the
250 Maglemose and Ertebølle cultures in Scandinavia, the Cardial in the Mediterranean, the Körös and
251 Linear Pottery (LBK) in SE and Central Europe, and many archaeological cultures in Ukraine,
252 western Russia, and the trans-Ural (e.g. Veretye, Lyalovo, Volosovo, Kitoi). Our sampling was
253 particularly dense in Denmark from where we present a detailed and continuous sequence of 100
254 genomes spanning from the early Mesolithic to the Bronze Age. Dense sample sequences were also
255 obtained from Ukraine, Western Russia, and the trans-Ural, spanning from the Early Mesolithic
256 through the Neolithic, up to c. 5,000 BP.

Edit: Their ISOGG labelling is really lazy, though: For instance, they have almost all the well-documented Driffield guys listed as: R1b1a1b1a1, which is P310/L52.
P312 is R1b1a1b1a1a2 and U106 is R1b1a1b1a1a1

Samples that need more detail (if coverage allows):

NEO870	Toftum mose	Denmark	NorthernEurope	Denmark_Neolithic	55.895	9.987	thisStudy	4,404	4,105	4,255	direct	XY	R1b	R1b1a1b1a1	0.58	0.993	0	1.2.1.3.1.2	Scandivia_4200BP_3200BP	Eurasia_5000BP_200BP	PostNeol_EuropeW
NEO872	Toftum mose	Denmark	NorthernEurope	Denmark_Neolithic	55.895	9.987	thisStudy	4,140	3,891	4,016	direct	XY	R1b	R1b1a1b1a1	0.08	0.980	lowcov				
NEO92	Barhøj (Strøby Egede)	Denmark	NorthernEurope	Denmark_Neolithic	55.403	12.272	thisStudy	4,348	4,088	4,218	direct	XY	R1b	R1b1a1b1a1	0.61	0.993	0	1.2.1.3.1.1	Scandivia_4200BP_3200BP	Eurasia_5000BP_200BP	PostNeol_EuropeW
NEO735	Borreby	Denmark	NorthernEurope	Denmark_Neolithic	55.225	11.303	thisStudy	4,138	3,852	3,995	direct	XY	R1b	R1b1a1b1a1	0.67	0.994	0	1.2.1.3.1.4	Scandivia_4200BP_3200BP	Eurasia_5000BP_200BP	PostNeol_EuropeW
NEO737	Borreby	Denmark	NorthernEurope	Denmark_Neolithic	55.225	11.303	thisStudy	4,236	3,985	4,111	direct	XY	R1b	R1b1a1b1a1	0.24	0.988	0	1.2.1.2.2.1.2.2	Europe_4500BP_2000BP	Eurasia_5000BP_200BP	PostNeol_EuropeW
NEO860	Mosede Mose	Denmark	NorthernEurope	Denmark_Neolithic	55.576	12.275	thisStudy	3,835	3,498	3,667	direct	XY	R1b	R1b1a1b1a1	0.20	0.988	0	1.2.1.3.1.1	Scandivia_4200BP_3200BP	Eurasia_5000BP_200BP	PostNeol_EuropeWNEO735	Borreby	Denmark	NorthernEurope	Denmark_Neolithic	55.225	11.303	thisStudy	4,138	3,852	3,995	direct	XY	R1b	R1b1a1b1a1	0.67	0.994	0	1.2.1.3.1.4	Scandivia_4200BP_3200BP	Eurasia_5000BP_200BP	PostNeol_EuropeW

Edit: Found more from Sweden:

NEO52	Vattenledningen	Sweden	NorthernEurope	Sweden_Neolithic	55.503	13.009	thisStudy	3,885	3,646	3,766	direct	XY	R1b	R1b1a1b1a1	0.11	0.983	0	1.2.1.2.2.1.1.4	Europe_4500BP_2000BP	Eurasia_5000BP_200BP	PostNeol_EuropeW

[Max_H]

Population Genomics of Stone Age Eurasia

Allenthoft & Willerslev et. al. 2022, Biorxiv.

Abstract:

Interesting paper, but images of low quality. However, how accurate is Extended Data Fig. 3 of IBD sharing? Some of the samples share unlikely connections, for example Ustishim with younger populations.

In addition, I do not believe there is a peak of EHG-like ancestry in Mongolia.

[etrusco]

Am I wrong or Middle Don hg are a combination of a Ukraine hg population admixed with a CHG like population.
That is evident in fig.2
As I said many times now is finally confirmed : Dneper Don foragers=PIE
Evident their cultural affiliation with the Mariupol cultural horizon

[RCO]

Important paper

I think that's a very interesting discovery

We demonstrate that this “steppe” ancestry (Steppe_5000BP_4300BP) can be modelled as a 428 mixture of ~65% ancestry related to herein reported hunter-gatherer genomes from the Middle Don 429 River region (MiddleDon_7500BP) and ~35% ancestry related to hunter-gatherers from Caucasus 430 (Caucasus_13000BP_10000BP) (Extended Data Fig. 4). Thus, Middle Don hunter-gatherers, who 431 already carry ancestry related to Caucasus hunter-gatherers (Fig. 2), serve as a hitherto unknown 432 proximal source for the majority ancestry contribution into Yamnaya genomes. The individuals in 433 question derive from the burial ground Golubaya Krinitsa (Supplementary Note 3). Material culture 434 and burial practices at this site are similar to the Mariupol-type graves, which are widely found in 435 neighbouring regions of Ukraine, for instance along the Dnepr River. They belong to the group of 436 complex pottery-using hunter-gatherers mentioned above, but the genetic composition at Golubaya 437 Krinitsa is different from the remaining Ukrainian sites (Fig 2A, Extended Data Fig. 4).

(...) An influx of southern European hunter-gatherer
381 related ancestry in Ukrainian individuals after the Mesolithic (Fig. 3) suggests a similar eastwards
382 expansion in south-eastern Europe17. Interestingly, two herein reported ~7,300-year-old imputed
383 genomes from the Middle Don River region in the Pontic-Caspian steppe (Golubaya Krinitsa,
384 NEO113 & NEO212) derive ~20-30% of their ancestry from a source cluster of hunter-gatherers
385 from the Caucasus (Caucasus_13000BP_10000BP) (Fig. 3). Additional lower coverage (non
386 imputed) genomes from the same site project in the same PCA space (Fig. 1D), shifted away from
387 the European hunter-gatherer cline towards Iran and the Caucasus. Our results thus document
388 genetic contact between populations from the Caucasus and the Steppe region as early as 7,300
389 years ago, providing documentation of continuous admixture prior to the advent of later nomadic
390 Steppe cultures, in contrast to recent hypotheses, and also further to the west than previously
391 reported17,37 391 .

Supplementary Tables[supplements/490594_file04.xlsx

NEO113 Golubaya Krinitsa Russia CentralEasternEurope Russia_Neolithic_MiddleDon 50.073 39.878 thisStudy 7,423 7,172 7,298 direct XY R1a R1a 0.11 0.983 0 4.1.3.2.2 MiddleDon_7500BP Europe_15000BP_4000BP HG_EuropeE

NEO212 Golubaya Krinitsa Russia CentralEasternEurope Russia_Neolithic_MiddleDon 50.072 39.894 thisStudy 7,475 7,311 7,393 direct XY I2a I2a1b1a2 3.18 0.996 0 4.1.3.1.1.1 MiddleDon_7500BP Europe_15000BP_4000BP HG_EuropeE

And they also found one J1 here:
NEO163 Vasilyevskiy kordon 17 Russia CentralEasternEurope Russia_Neolithic 52.9 40.03 thisStudy 5,721 5,486 5,604 direct XY J1 J1 0.23 0.990 0 4.1.1.2.1.2 DonRiver_5800BP_5300BP Europe_15000BP_4000BP HG_EuropeE

[Gentica277282]

It will be very interesting to have these samples added to the database

[Gentica277282]

Hmmm I think it is the original Mesolithic Kotias Klde sample. It has the exact same mtDNA clade ad KK1. He should be listed as J2a. Kind of weird how they are calling him Neolithic in this paper as well.

Do we know what subclade his under

[Ajeje Brazorf]

Group label	Sample	Site	Date	Y-DNA	mtDNA	Latitude and Longitude
Georgia_UpperPaleolithic	NEO283	Kotias Klde	-23685		U4'9	42.22, 43.32
Ukraine_Mesolithic	NEO550	Vasilevka-I	-9086		H2a2a1	48.2, 35.216
Ukraine_Mesolithic	NEO527	Voloshskoe	-8974		U2e1	48.305, 35.136
Russia_Mesolithic	NEO202	Peschanitsa	-8903	R1a	U4a1	61.272, 38.91
Ukraine_Mesolithic	NEO545	Vasilevka-I	-8879	I	H2a	48.2, 35.216
Ukraine_Mesolithic	NEO494	Vasilevka-I	-8791	R1	U5a1d	48.2, 35.216
Ukraine_Mesolithic	NEO496	Vasilevka-I	-8705	GHIJK	H2a2a1	48.2, 35.216
Ukraine_Mesolithic	NEO492	Vasilevka-I	-8705	I2a1b	U2e	48.2, 35.216
Ukraine_Mesolithic	NEO497	Vasilevka-I	-8705	R1a	U4	48.2, 35.216
Ukraine_Mesolithic	NEO501	Vasilevka-I	-8672	Q	U5a2	48.2, 35.216
Ukraine_Mesolithic	NEO549	Vasilevka-I	-8615	I	H2a2a1	48.2, 35.216
Denmark_Mesolithic	NEO254	Koelbjerg	-8515	I2a1b	U5a2c	55.404, 10.133
Russia_Mesolithic	NEO539	Minino	-8107		U4	59.7, 39.5
Russia_Mesolithic	NEO100	Ural River Beach	-7977	Q1b	H2a2a1	51.566, 53.683
Sweden_Mesolithic	NEO18	Hanaskede	-7925	I2a1a	U5a1	58.391, 13.558
Serbia_Mesolithic	NEO669	Lepenski Vir	-7850		H13	44.5555278, 22.02742
Georgia_Mesolithic	NEO281	Kotias Klde	-7773	J2b	H13c	42.22, 43.32
Sweden_Mesolithic	NEO27	Bredgården	-7743	I2a1a	U5a2	57.742, 13.403
Russia_Mesolithic	NEO536	Minino	-7618	R1b	U4	59.7, 39.5
Spain_Mesolithic	NEO694	Santa Maira	-7572	I2a1a	U2'3'4'7'8'9	38.73, -0.463
Denmark_Mesolithic	NEO13	Hedegaard (Bislev)	-7561	I	H2a2a1	56.907, 9.576
Iran_Neolithic	NEO817	Tepe Guran	-7442		H2a2a1	33.756, 47.097
Iran_Neolithic	NEO819	Tepe Guran	-7341		U7a	33.756, 47.097
Norway_Mesolithic	NEO17	Hummervikholmen	-7197	I2	U4	58.064, 7.743
Denmark_Mesolithic	NEO91	Strøby Grøftemark	-7184	I2a1a2	K1e	55.393, 12.307
Denmark_Mesolithic	NEO759	Køge Sønakke	-7090	I2	[email protected]	55.402, 12.367
Romania_Mesolithic	NEO672	Schela Cladovei	-6864		K1b2	44.635, 22.66
Iran_Neolithic	NEO816	Tepe Guran	-6750	J2a	R2	33.756, 47.097
Serbia_Mesolithic	NEO677	Vlasac	-6726	R1b1a	K1+16362	44.5355833, 22.0444444
Serbia_Mesolithic	NEO655	Vlasac	-6726		HV+16311	44.5355833, 22.0444444
Serbia_Mesolithic	NEO657	Vlasac	-6726		K1b1b1	44.5355833, 22.0444444
Romania_Mesolithic	NEO671	Schela Cladovei	-6709	R1b	U4a	44.635, 22.66
Russia_Siberia_Neolithic_Koshinskaya	NEO72	Mergen' 6	-6406	Q1b	R1b1	56.019, 69.31
Russia_Neolithic	NEO555	Karavaikha	-6327	R1b1a1	T2a1b1	60.414, 38.932
Spain_Mesolithic	NEO646	El Mazo	-6324		U5b1b1	43.404, -4.707
Ukraine_Neolithic	NEO521	Igren	-6320		U5b2b	48.396, 35.161
Russia_Neolithic	NEO559	Karavaikha	-6318	R1b1a1	U5a1	60.414, 38.932
Russia_Mesolithic	NEO87	Zamostje 2	-6254		U5a1	56.666, 38.016
Russia_Siberia_Neolithic	NEO116	Pad’ Tokui	-6237		C4a1a1	49.8, 108.8
Russia_Neolithic	NEO561	Karavaikha	-6234		U5a1	60.414, 38.932
Denmark_Mesolithic	NEO123	Orehoved Sejlrende	-6228		U5b1b	54.96, 11.85
Denmark_Mesolithic	NEO19	Røntesten	-6222	I2a1b	U5a2b	56.27, 10.472
Denmark_Mesolithic	NEO122	Orehoved Sejlrende	-6206		U5a2b	54.96, 11.85
Russia_Neolithic	NEO558	Karavaikha	-6135	R1a	K1+16362	60.414, 38.932
Russia_Neolithic	NEO554	Pogostishche I	-6115		U5a2d	60.45, 38.633
Russia_Mesolithic	NEO537	Minino	-6102	R1b1a	U4a	59.7, 39.5
Serbia_Mesolithic	NEO658	Lepenski Vir	-6065		N1a1a1a2	44.5355833, 22.0444444
Russia_Neolithic	NEO557	Karavaikha	-5962	R1b1a1	U4	60.414, 38.932
Russia_Neolithic	NEO560	Karavaikha	-5955	R1b1a1	U5a1	60.414, 38.932
Spain_Mesolithic	NEO938	El Mazo	-5928		U5a2a	43.404, -4.707
Russia_Mesolithic	NEO88	Zamostje 2	-5908	R1b1a1	U5a1c	56.666, 38.016
Denmark_Mesolithic	NEO600	Vedbaek boldbaner	-5865	I2	K1e	55.848, 12.56
Hungary_Neolithic_Koros	NEO130	Dezsk	-5853		V1	46.251, 20.259
Russia_Siberia_Neolithic_Koshinskaya	NEO73	Mergen' 6	-5731		U2e3	56.019, 69.31
Russia_Siberia_Neolithic	NEO117	Zhindo	-5706	N1a2	C4a1a+195	49.9, 108
Armenia_Neolithic	NEO110	Aknashen	-5644		T1a	40.101, 44.294
Hungary_Neolithic_Koros	NEO137	Hödmezövasarhely Kotac	-5642		H5b	46.418, 20.33
Russia_Siberia_Neolithic	NEO115	Zhindo	-5600	N1a2	[email protected]	49.9, 108
Russia_Siberia_Neolithic	NEO922	Vengerovo-2	-5599	Q1b2	U2e3	55.707, 76.783
Spain_Mesolithic	NEO648	El Toral III	-5574	I2a1a	U5a2a	43.402, -4.707
Denmark_Mesolithic	NEO683	Tybrind Vig	-5571		U5a2b	55.395, 9.826
Denmark_Mesolithic	NEO589	Korsør Nor	-5525	I2a1a2	U5a2b	55.331, 11.145
Latvia_Mesolithic	NEO307	Zvejnieki	-5506	I2a1b1a2	U5a2d	57.775, 25.229
Italy_Neolithic	NEO834	Mora Cavorso	-5460	G2a2a	T2b	41.87, 13.283
Russia_Neolithic_MiddleDon	NEO212	Golubaya Krinitsa	-5443	I2a1b1a2	U4a2a	50.072, 39.894
Denmark_Mesolithic	NEO587	Kongemose	-5427	I2a1a2	U5b1	55.583, 11.551
Portugal_Neolithic_Cardial	NEO632	Gruta do Caldeirao	-5397	I2a1a	J2b1a	39.648, -8.415
Denmark_Mesolithic	NEO932	Tudse Hage	-5352		K1e	55.249, 11.228
Russia_Neolithic_MiddleDon	NEO113	Golubaya Krinitsa	-5348	R1a	U2e1a	50.073, 39.878
Russia_Neolithic_MiddleDon	NEO209	Golubaya Krinitsa	-5346	R1a	U5b2	50.072, 39.894
Italy_Neolithic	NEO695	Maddalena di Muccia	-5344		J1c3j	43.075, 13.059
Ukraine_Neolithic	NEO270	Mamaj Gora	-5343	I2a1b	U4d	47.433, 34.268
Ukraine_Neolithic	NEO551	Vovnigi I	-5336		U5a2+16362	48.122, 35.096
Ukraine_Neolithic	NEO268	Mamaj Gora	-5295	R1b	U5a2+16294	47.433, 34.268
Russia_Neolithic_MiddleDon	NEO204	Golubaya Krinitsa	-5278	I2a1b	U4	50.072, 39.894
Russia_Siberia_Neolithic_Kitoi	NEO199	Fofonovo	-5219	Q1b1	R11	52.2, 107
Russia_Siberia_Neolithic_Kitoi	NEO841	Sosnovyiy Myis	-5206		A+152+16362+16189	58.752, 102.634
Czech_Neolithic_LBK	NEO128	Vedrovice	-5191	G	K1a	49.021, 16.38
Portugal_Neolithic_Cardial	NEO631	Gruta do Caldeirao	-5185	I2a1a	T2b3+151	39.648, -8.415
Denmark_Mesolithic	NEO748	Henriksholm-Bøgebakken (Vedbæk)	-5179	I2	U5b1d2	55.853, 12.559
Denmark_Mesolithic	NEO814	Bodal K	-5165		U4a2	55.586, 11.538
Russia_Neolithic_MiddleDon	NEO210	Golubaya Krinitsa	-5158	R1	U5a2b	50.072, 39.894
Russia_Neolithic_MiddleDon	NEO207	Golubaya Krinitsa	-5147		U5a1c	50.072, 39.894
Ukraine_Neolithic	NEO516	Igren	-5146	Q1b	U5a1i	48.396, 35.161
Denmark_Mesolithic	NEO749	Henriksholm-Bøgebakken (Vedbæk)	-5122	I2a1a2	U5b1d2	55.853, 12.559
Russia_Siberia_Neolithic_Kitoi	NEO201	Fofonovo	-5114	Q1b1	C4	52.2, 107
Russia_Siberia_Neolithic_Kitoi	NEO843	Sosnovyiy Myis	-5110		R11	58.752, 102.634
Russia_Siberia_Neolithic_Sredneirtyshskaya	NEO75	Omskaya Stoyanka II	-5108	Q1a2	T2d	54.971, 73.366
Russia_Siberia_Neolithic	NEO63	Itkul	-5107	Q1a	A+152+16362	52.713, 84.692
Ukraine_Neolithic	NEO498	Vovnigi II	-5106	R1b1b	U4a1	48.132, 35.081
Russia_Siberia_Neolithic	NEO67	Ust'-Isha	-5098	C2	U5a2	52.18, 85.965
Denmark_Mesolithic	NEO791	Korsör Nor	-5083	I2	U5b1	55.331, 11.145
Denmark_Mesolithic	NEO586	Koed	-5080	I2	U5a2c	56.371, 10.573
Russia_Neolithic	NEO556	Karavaikha	-5080		U4a1	60.414, 38.932
Russia_Neolithic_Lyalovo	NEO185	Sakhtish IIa	-5078	Q1b1	U4a1	56.781, 40.447
Russia_Siberia_Neolithic_Kitoi	NEO200	Fofonovo	-5057	Q1b1	C4	52.2, 107
Ukraine_Neolithic	NEO304	Volnensky	-5056	I2a1b	U5a2d	47.95, 35.39
Denmark_Mesolithic	NEO746	Henriksholm-Bøgebakken (Vedbæk)	-5036	I	U4a2	55.853, 12.559
Denmark_Mesolithic	NEO583	Koed	-5030		U5a2b	56.371, 10.573
Hungary_Neolithic_Tisza	NEO149	Gorzsa Cukormajor	-5028	I	N1a1a1a	46.369, 20.424
Ukraine_Neolithic	NEO553	Vovnigi I	-5027		U5a	48.122, 35.096
Ukraine_Neolithic	NEO524	Vovnigi II	-5026	I2a1b	U5a1j	48.132, 35.081
Denmark_Mesolithic	NEO822	Dragsholm	-5023		U5a2b	55.768, 11.388
Ukraine_Neolithic	NEO552	Vovnigi I	-5020	I2a1b	U2e1	48.122, 35.096
Russia_Siberia_Neolithic_Sredneirtyshskaya	NEO78	Omskaya Stoyanka II	-5003	Q1a2	U4a1	54.971, 73.366
Russia_Neolithic_Lyalovo	NEO186	Sakhtish IIa	-4969	R1b1a1	U5a1d	56.781, 40.447
Sweden_Mesolithic	NEO679	Skateholm I	-4957		U5b2a1a	55.386, 13.481
Denmark_Mesolithic	NEO930	Fannerup F	-4942	I2	U5b2b1a	56.4, 10.714
Ukraine_Neolithic	NEO507	Vovnigi II	-4937	R1b	U5b2	48.132, 35.081
Russia_Neolithic_Lyalovo	NEO192	Sakhtish II	-4892		K1b2	56.78, 40.447
Ukraine_Neolithic	NEO528	Vovnigi I	-4892	I2a1b	T2	48.122, 35.096
Ukraine_Neolithic	NEO302	Volnensky	-4889	I2a1b1a2	U4a	47.95, 35.39
Denmark_Mesolithic	NEO733	Dragsholm	-4869		U5b2b	55.768, 11.388
Denmark_Mesolithic	NEO732	Dragsholm	-4862		U5b2b	55.768, 11.388
Denmark_Mesolithic	NEO745	Henriksholm-Bøgebakken (Vedbæk)	-4851	I2	U5b1d2	55.853, 12.559
Russia_Siberia_Neolithic	NEO911	Vengerovo-2	-4830	Q1b2	C5c	55.707, 76.783
Denmark_Mesolithic	NEO856	Nederst	-4828		U5a2b	56.365, 10.643
Russia_Neolithic	NEO172	Ksizovo 6	-4824	R1b	U5a2b2	52.278, 38.955
Hungary_Neolithic_Tisza	NEO145	Gorzsa Cukormajor	-4799		T2c1d1	46.369, 20.424
Ukraine_Neolithic	NEO529	Vovnigi I	-4794		U5a1	48.122, 35.096
Denmark_Mesolithic	NEO747	Henriksholm-Bøgebakken (Vedbæk)	-4778	I2a1b	R1b1	55.853, 12.559
Hungary_Neolithic_Tisza	NEO147	Gorzsa Cukormajor	-4774		J1c6	46.369, 20.424
Hungary_Neolithic_Tisza	NEO140	Gorzsa Cukormajor	-4766	H	T2b23	46.369, 20.424
Ukraine_Neolithic	NEO503	Vovnigi II	-4756		U5a2d	48.132, 35.081
Russia_Neolithic	NEO162	Vasilyevskiy kordon 17	-4731		U5a1d1	52.9, 40.03
Ukraine_Neolithic	NEO300	Volnensky	-4728		U5b2	47.95, 35.39
Hungary_Neolithic_Tisza	NEO143	Gorzsa Cukormajor	-4706	J2a	T2c1d1	46.369, 20.424
Hungary_Neolithic_Tisza	NEO142	Gorzsa Cukormajor	-4675		U1a1a	46.369, 20.424
Ukraine_Neolithic	NEO514	Vovnigi II	-4673	I2	H2a	48.132, 35.081
Ukraine_Neolithic	NEO502	Vovnigi II	-4666	I2a1b	U5b1	48.132, 35.081
Hungary_Neolithic_Tisza	NEO148	Gorzsa Cukormajor	-4662		U5b2c	46.369, 20.424
Ukraine_Neolithic	NEO305	Vasilevsky	-4661	I2a1b1a2	U4a	48.305, 35.139
Denmark_Mesolithic	NEO1	Holmegård-Djursland	-4644	I2	U5a'b	56.251, 10.755
Turkmenistan_Neolithic	NEO310	Monjukli-Depe	-4600	L1a2	H14	36.848, 60.418
France_Neolithic_Cardial	NEO812	Grotte du Gazel	-4595		H1	43.324, 2.42
France_Neolithic_Cardial	NEO813	Grotte du Gazel	-4591		H1	43.324, 2.42
Ukraine_Neolithic	NEO262	Lysa Gora	-4563	R1b	[email protected]	47.451, 35.239
Denmark_Mesolithic	NEO941	Havnø	-4424		[email protected]	56.709, 10.171
Denmark_Mesolithic	NEO570	Fannerup E	-4422		U5b2c	56.401, 10.718
Russia_Neolithic	NEO173	Ksizovo 6	-4393		U5a2b2	52.278, 38.955
Denmark_Mesolithic	NEO751	Bjørnsholm	-4365	I2	U5b2b1a	56.874, 9.215
Denmark_Mesolithic	NEO852	Norsminde	-4356		U5b1	56.027, 10.257
Ukraine_Neolithic	NEO508	Vovnigi II	-4354		U4a1	48.132, 35.081
Denmark_Mesolithic	NEO855	Fannerup D	-4349	I2a1b1a2	[email protected]	56.401, 10.715
Denmark_Mesolithic	NEO568	Ertebølle	-4347	I2	U5b1	56.812, 9.179
Denmark_Mesolithic	NEO569	Ertebølle	-4195	I2	U5b2a1a	56.812, 9.179
Denmark_Mesolithic	NEO3	Vængesø II	-4142	I	U5a	56.146, 10.552
Denmark_Mesolithic	NEO598	Sølager	-4118		R1b1	55.948, 11.897
Ukraine_Neolithic	NEO522	Voloshskoe	-4105	I2a1b	U5a1	48.305, 35.136
Russia_Siberia_Neolithic	NEO64	Itkul	-4100	C2b	R1b1	52.713, 84.692
Denmark_Mesolithic	NEO853	Langø Skaldynge	-4095		R1b1	55.545, 10.622
Russia_Siberia_Neolithic	NEO292	Dolgoe Ozero	-4086	Q1b1	A+152+16362+16189	56.2, 95.7
Russia_Neolithic_Volosovo	NEO181	Sakhtish IIa	-4084	Q1b2a	U2e1e	56.781, 40.447
Denmark_Mesolithic	NEO960	Ravnsbjerggård II	-4013	I2a1a	U5b2a2	55.581, 11.578
Russia_Neolithic_Volosovo	NEO180	Sakhtish IIa	-3989	R1b1a1	U5b2c1	56.781, 40.447
Denmark_Mesolithic	NEO645	Rødhals	-3886	I2	K1e	55.914, 11.091
Russia_Neolithic	NEO171	Vasilyevskiy kordon 17	-3885		[email protected]	52.9, 40.03
Russia_Siberia_Neolithic	NEO309	Protoka	-3874	Q1b2	A10	56.77, 77.03
Sweden_Mesolithic	NEO260	Evensås	-3872	I2	R1b1	58.246, 11.492
Russia_Siberia_Neolithic	NEO923	Vengerovo-2	-3864		A10	55.707, 76.783
Denmark_Neolithic	NEO601	Viksø Mose	-3857		K1a2b	55.759, 12.238
Denmark_Mesolithic	NEO962	Dragsholm	-3836	I	U5b2b	55.768, 11.388
Russia_Siberia_Neolithic	NEO102	Afontova Gora	-3831	Q	U2e1h	56.016, 92.866
Russia_Neolithic_Volosovo	NEO158	Sakhtish II	-3804	R1b1a	U2e1e	56.78, 40.447
Russia_Neolithic_Volosovo	NEO195	Sakhtish IIa	-3797	I2a1a	U5a2d	56.781, 40.447
Russia_Siberia_Neolithic	NEO916	Vengerovo-2	-3777	K2b	H2a2a1	55.707, 76.783
Russia_Siberia_Neolithic	NEO915	Vengerovo-2	-3777	Q1	C4+152	55.707, 76.783
Denmark_Neolithic	NEO891	Roskilde Fjord (syd for Jyllinge)	-3764		H	55.73, 12.1
Russia_Neolithic	NEO164	Vasilyevskiy kordon 17	-3719	R1a	U5a1d1	52.9, 40.03
Denmark_Neolithic	NEO790	Tysmose	-3718	I2a1a2	HV16	55.711, 12.274
Russia_Siberia_Neolithic	NEO83	Borovjanka XVII	-3714	R1b1a1	U4a	55.733, 74.285
Russia_Neolithic	NEO166	Vasilyevskiy kordon 17	-3713	R1b1a1	U5a1f1	52.9, 40.03
Russia_Neolithic	NEO167	Vasilyevskiy kordon 17	-3706		U2e1e	52.9, 40.03
Russia_Siberia_Neolithic	NEO907	Vengerovo-2	-3699	C2	A8a	55.707, 76.783
Russia_Neolithic	NEO163	Vasilyevskiy kordon 17	-3654	J1	U5a2	52.9, 40.03
Russia_Neolithic_Volosovo	NEO189	Sakhtish IIa	-3654	R1a	U5a2d	56.781, 40.447
Russia_Neolithic_Volosovo	NEO194	Sakhtish IIa	-3647	R1b1a1	U4b1b1	56.781, 40.447
Romania_Neolithic	NEO674	Baile Herculane	-3640	I2a1b	U8b1b1	44.896, 22.428
Sweden_Neolithic_FBC	NEO46	Ängdala	-3614		T2b	55.588, 13.113
Russia_Neolithic	NEO170	Vasilyevskiy kordon 17	-3613		U2e1e	52.9, 40.03
Denmark_Neolithic	NEO29	Lohals	-3609		H	55.134, 10.903
Denmark_Neolithic	NEO595	Pandebjerg	-3595	I2a1a2	J1c	55.789, 11.292
Denmark_Neolithic	NEO23	Jorløse mose	-3586	I2a1a2	U5b1b2	55.601, 11.312
Russia_Siberia_Neolithic	NEO912	Vengerovo-2	-3586		C4+152	55.707, 76.783
Denmark_Neolithic	NEO564	Bygholm Nørremark	-3585	I2a1a	J1c3j	55.875, 9.812
Denmark_Neolithic	NEO571	Grøfte	-3585		H2a2a	55.404, 11.497
Ukraine_Neolithic	NEO265	Lysa Gora	-3585	I2a1b	U5a2+16362	47.451, 35.239
Denmark_Neolithic	NEO753	Sigersdal Mose 2	-3583		J1c	55.772, 12.213
Denmark_Neolithic	NEO41	Rude	-3581	I	J2a1a1a	55.994, 10.248
Denmark_Neolithic	NEO942	Tissøe	-3545		T2b	55.575, 11.294
Russia_Neolithic_Volosovo	NEO179	Sakhtish II	-3515		U5a2d	56.78, 40.447
Russia_Siberia_Neolithic	NEO917	Vengerovo-2	-3512	Q1a2	C4	55.707, 76.783
Britain_Neolithic	NEO847	Hetty Peglers Tump	-3511	I2a1b1a1a	X2b4	51.697, -2.304
Denmark_Neolithic	NEO28	Salpetermosen syd 8	-3511	I2a1a2	T2b11	55.91, 12.307
Denmark_Neolithic	NEO886	Dalmosegaard	-3508	I2a1a2	H26	54.989, 12.419
Russia_Siberia_Neolithic	NEO918	Vengerovo-2	-3508		U2e2a	55.707, 76.783
Denmark_Neolithic	NEO866	Lundby-Falster	-3507	R1b1a	K1a2b	54.865, 11.841
Russia_Neolithic_Volosovo	NEO184	Sakhtish 8	-3507		U5a2d	56.776, 40.443
Russia_Neolithic_Volosovo	NEO183	Sakhtish IIa	-3506	R1b	[email protected]	56.781, 40.447
Russia_Neolithic	NEO168	Vasilyevskiy kordon 17	-3503		[email protected]	52.9, 40.03
Denmark_Neolithic	NEO757	Sejerby (Sejerø)	-3502		T2b	55.902, 11.118
Russia_Neolithic_Volosovo	NEO193	Sakhtish IIa	-3501	R1b	U2e1e	56.781, 40.447
Denmark_Neolithic	NEO896	Mandemarke	-3494		H4a1a1a	54.969, 12.494
Denmark_Neolithic	NEO945	Læsten Mose	-3494		H1b1	56.485, 9.827
Russia_Neolithic	NEO687	Kumyshanskaya Cave	-3493	R1a	U5a1d2	57.58, 58.2
Denmark_Neolithic	NEO933	Sludegård Sømose	-3488	I2	V	55.253, 10.746
Italy_Neolithic	NEO830	Fontenoce	-3443		I1c1a	43.383, 13.55
Denmark_Neolithic	NEO43	Rude	-3442	I	J2a1a1a	55.994, 10.248
Russia_Neolithic_Volosovo	NEO188	Sakhtish IIa	-3440	I2a1b	U5a1f1	56.781, 40.447
Denmark_Neolithic	NEO888	Elkenøre	-3439		H5	54.732, 11.986
Russia_Neolithic_Volosovo	NEO182	Sakhtish IIa	-3436	R1b	[email protected]	56.781, 40.447
Denmark_Neolithic	NEO744	Vig Femhøve	-3413	Q1b2	K1a2a	55.856, 11.59
Denmark_Neolithic	NEO795	Porsmose	-3380	I2a1a2	H24	55.276, 11.875
Russia_Neolithic	NEO174	Ksizovo 6	-3357		W6a	52.278, 38.955
Denmark_Neolithic	NEO702	Jørlundegård	-3321	I	K1a1a2	55.828, 12.166
Russia_Neolithic_Volosovo	NEO178	Sakhtish II	-3318	I2a1a	U5a1d	56.78, 40.447
Russia_Neolithic	NEO160	Vasilyevskiy kordon 17	-3313		U5a1d1	52.9, 40.03
Denmark_Neolithic	NEO935	Vibygårds Mose	-3298	I2a1a2	H	55.564, 12.016
Russia_Neolithic_Volosovo	NEO197	Sakhtish IIa	-3295	I2a1a	U5a2d	56.781, 40.447
Denmark_Neolithic	NEO7	Sigersdal Mose	-3290	CT	H	55.761, 12.258
Denmark_Neolithic	NEO597	Storelyng (Øgårde boat III)	-3264	I2a1a2	U5b1b2	55.587, 11.574
Denmark_Neolithic	NEO865	Lundby-Falster	-3234	I2	J1c3g	54.865, 11.841
Denmark_Neolithic	NEO594	Neverkær mose	-3231	I2a1b1a2	K1a+195	55.391, 10.078
Denmark_Neolithic	NEO961	Avlebjerg (Strøby)	-3199		K1b1a1	55.368, 12.287
Denmark_Neolithic	NEO599	Vanløse mose	-3191	Q1b	K2b1a	55.553, 11.684
Denmark_Neolithic	NEO602	Storelyng (Østrup Homo II)	-3189	I2	U5b2a1b	55.589, 11.576
Denmark_Neolithic	NEO33	Vittrup	-3189	I2a1a	H2a	57.391, 9.761
Denmark_Neolithic	NEO566	Døjringe	-3185	I2a1a	U5b3	55.484, 11.579
Britain_Neolithic	NEO627	Banks tomb	-3182	I2a1a2	U5b1c	58.734, -2.937
Poland_Neolithic_FBC	NEO641	Slonowice	-3182		H7d	50.267, 20.45
Sweden_Neolithic	NEO36	Hindby mosse	-3147		K2b1a	55.565, 13.036
Britain_Neolithic	NEO626	Banks tomb	-3132	I2a1a2	H1	58.734, -2.937
Denmark_Neolithic	NEO898	Svinninge Vejle	-3131	I2a1a2	K1a3a	55.736, 11.425
Britain_Neolithic	NEO625	Banks tomb	-3130	I2a1a2	U5b1c	58.734, -2.937
Sweden_Neolithic	NEO39	Hindby mosse	-3124	I2a1a2	H49	55.565, 13.036
Sweden_Neolithic	NEO38	Hindby mosse	-3121		K1a3	55.565, 13.036
Denmark_Neolithic	NEO25	Kainsbakke II	-3116		K1a+195	56.433, 10.789
Sweden_Neolithic	NEO259	Frälsegården	-3010	I2a1a	K2a	58.164, 13.454
Denmark_Neolithic	NEO925	Myrebjerg mose	-2997		W1	54.771, 10.675
Russia_Neolithic_Volosovo	NEO187	Sakhtish IIa	-2990		U2e1e	56.781, 40.447
Poland_Neolithic_FBC	NEO640	Slonowice	-2950	G2a2b2a	H	50.267, 20.45
Russia_Siberia_BronzeAge_Okunevo	NEO68	Okunevo 5 and 7	-2949		D4j	56.438, 74.907
Britain_Neolithic	NEO630	Banks tomb	-2947	I2	H67	58.734, -2.937
Britain_Neolithic	NEO624	Banks tomb	-2947		K1b1a1	58.734, -2.937
Britain_Neolithic	NEO717	Banks tomb	-2900	I2a1a2	K1a+195	58.734, -2.937
Russia_Siberia_BronzeAge	NEO65	Kostenkova Isbushka	-2854	C2	A8a1	52.724, 84.683
Russia_Siberia_BronzeAge	NEO80	Borovjanka XVII	-2842		C	55.733, 74.285
Italy_Neolithic	NEO828	Gaudo	-2796	J2a	N1a1a1	40.416, 15.066
Russia_Siberia_BronzeAge_Okunevo	NEO77	Okunevo 5 and 7	-2782	Q1b	C4+152	56.438, 74.907
Italy_BronzeAge	NEO823	Grotta Nisco	-2715	G2a2b	U1a1a	40.883, 16.733
Russia_Siberia_BronzeAge_Okunevo	NEO79	Okunevo 5 and 7	-2681		U2e1'2'3	56.438, 74.907
Denmark_Neolithic	NEO943	Stenderup Hage	-2667	I2a1b1a1b	H+152	55.46, 9.694
Denmark_Neolithic	NEO580	Klokkehøj	-2662		H2a2a1	55.108, 10.111
Russia_Neolithic	NEO175	Ksizovo 6	-2657	I2a1b	U2e1'2'3	52.278, 38.955
France_Neolithic	NEO121	Grotte Mandrin	-2577	I2a1a1a1a1	U4a2f	44.466, 4.766
Estonia_Neolithic_CWC	NEO306	Sope	-2548		R1b1	59.408, 27.026
Denmark_Neolithic	NEO792	Næs	-2546	I2a1a2	U2e2a1a2	54.87, 12.116
Kazakhstan_BronzeAge	NEO57	Biestamak	-2546		A10	52.166, 64.533
Spain_Neolithic	NEO721	Camino de las Yeseras	-2545		U5b3	40.439, -3.501
Sweden_Neolithic	NEO44	Dösemarken	-2436		J1c2	55.564, 12.946
Denmark_Neolithic	NEO876	Toftum mose	-2435	R1a	T2a1a	55.895, 9.987
France_Neolithic	NEO119	Grotte Mandrin	-2432		U5b3	44.466, 4.766
Sweden_Neolithic	NEO51	Kastanjegården	-2391		U4d1	55.555, 13.023
Portugal_Neolithic	NEO603	Sao Paulo 2	-2387		[email protected]	38.676, -9.164
Portugal_Neolithic	NEO609	Sao Paulo 2	-2381	I2a1a	K1a1	38.676, -9.164
Denmark_Neolithic	NEO870	Toftum mose	-2305	R1b1a1b1a1	T2b2	55.895, 9.987
Denmark_Neolithic	NEO92	Barhøj (Strøby Egede)	-2268	R1b1a1b1a1	T2b	55.403, 12.272
Spain_BronzeAge	NEO649	El Toral III	-2166	R1b1a1b1a1	H1+16355	43.402, -4.707
Denmark_Neolithic	NEO738	Kolind	-2161	I2a1b1a2b	J1c3g	56.362, 10.593
Denmark_Neolithic	NEO737	Borreby	-2161	R1b1a1b1a1	U4b1a1a1	55.225, 11.303
Denmark_Neolithic	NEO861	Mosede Mose	-2161		H6a1a	55.576, 12.275
Spain_BronzeAge	NEO650	El Toral III	-2160	R1b1a1b1a1	J1c1	43.402, -4.707
Russia_Siberia_BronzeAge	NEO921	Vengerovo-2	-2148		R1b1	55.707, 76.783
Russia_Siberia_BronzeAge	NEO76	Ostrov 2	-2117		U4a2	56.933, 66.467
Ukraine_BronzeAge	NEO509	Igren	-2114	R1b1a1b1b	U5a1j	48.396, 35.161
Denmark_Neolithic	NEO878	Kyndeløse	-2088		U4a2	55.703, 11.855
Denmark_Neolithic	NEO872	Toftum mose	-2066	R1b1a1b1a1	K2a5	55.895, 9.987
Denmark_Neolithic	NEO735	Borreby	-2045	R1b1a1b1a1	U4c1	55.225, 11.303
Denmark_Neolithic	NEO875	Toftum mose	-2036	I1	U5b3b	55.895, 9.987
Sweden_Neolithic	NEO220	Falköping 5	-2036	I1	H13a1a	58.166, 13.573
Denmark_Neolithic	NEO739	Kolind	-2021	I2a1b1a2	N1a1a1a1	56.362, 10.593
Sweden_Neolithic	NEO223	Falköping 5	-2018	I1	W1b1	58.166, 13.573
Sweden_Neolithic	NEO227	Falköping 5	-1974	I1	H1a3	58.166, 13.573
Sweden_Neolithic	NEO224	Falköping 5	-1969		HV6	58.166, 13.573
Sweden_Neolithic	NEO261	Sillvik	-1945	I1	T2b	57.736, 11.744
Sweden_Neolithic	NEO228	Falköping 5	-1891	I1	H3h	58.166, 13.573
Denmark_Neolithic	NEO934	Gammellung	-1884		W1+119	54.876, 10.784
Kazakhstan_BronzeAge	NEO900	Sjauke	-1854	N1	U2e1	52.431, 76.838
Sweden_Neolithic	NEO52	Vattenledningen	-1816	R1b1a1b1a1	H3g4	55.503, 13.009
Sweden_Neolithic	NEO226	Falköping 5	-1811		I3d	58.166, 13.573
Sweden_Neolithic	NEO225	Falköping 5	-1798		U5b2a1a1	58.166, 13.573
Denmark_BronzeAge	NEO93	Strøby Ladeplads	-1788	I1	K1a1b2b	55.406, 12.285
Kazakhstan_BronzeAge	NEO904	Sjiderti 10	-1750		C4a1	51.651, 74.654
Sweden_Neolithic	NEO221	Falköping 5	-1748	R1a1a1b1a2b	H1a3	58.166, 13.573
Denmark_BronzeAge	NEO857	Lollikehuse	-1718	I1	H3v+16093	55.758, 12.007
Denmark_Neolithic	NEO860	Mosede Mose	-1717	R1b1a1b1a1	K1a1b2a1a	55.576, 12.275
Denmark_BronzeAge	NEO752	Madesø	-1699	R1b1a1b1a1a1b	U5a1i1	55.621, 11.331
Kazakhstan_BronzeAge	NEO899	Gregorievka 1	-1683		U2e1	52.629, 76.727
Russia_Siberia_BronzeAge	NEO910	Vengerovo-2	-1645		C4+152	55.707, 76.783
Denmark_BronzeAge	NEO815	Vasagård	-1524	I1	H3b6	55.055, 14.885
Spain_BronzeAge	NEO653	El Toral III	-1468		HV0d	43.402, -4.707
France_BronzeAge	NEO120	Grotte Mandrin	-1409		U5a2b3	44.466, 4.766
Denmark_BronzeAge	NEO563	Bybjerg	-1405	I1	H6a1a	55.76, 11.798
Denmark_BronzeAge	NEO951	Klæsterupholm Mose	-1356		W1	57.29, 10.033
Russia_BronzeAge	NEO61	Bol'shoy Oleni Ostrov	-1356		U5a2d	68.966, 33.083
Russia_BronzeAge	NEO60	Bol'shoy Oleni Ostrov	-1350		U5a1d2	68.966, 33.083
Denmark_BronzeAge	NEO590	Magleø	-1347	I1a2a1	K2a3	55.34, 11.161
Russia_Siberia_BronzeAge	NEO81	Borovjanka XVII	-1347	R1a	C5c	55.733, 74.285
Russia_BronzeAge	NEO62	Bol'shoy Oleni Ostrov	-1336		Z1a1a	68.966, 33.083
Denmark_BronzeAge	NEO946	Hove Å	-1195	R1b1a1b1a1a1b	U4a1a	55.708, 12.25
Italy_BronzeAge	NEO806	Grotta Delle Mura	-1063	J2b2a1a1a1a1a1b	N1a1a1	40.957, 17.29
Russia_Siberia_BronzeAge	NEO70	Bazaiha	-802	N1	F1b1b	55.956, 92.791
Ukraine_IronAge	NEO278	Kleshnya III	-793	R1a	U5b1a	48.966, 38.238
Russia_IronAge	NEO538	Minino	-300	N1a1a1a1a2	U5a1d2b	59.7, 39.5
Ukraine_IronAge	NEO518	Igren	-202	R1a1a1b2	U5a2d	48.396, 35.161
Kazakhstan_IronAge	NEO902	Sjauke 1	-36	Q1b	J1b1a1	52.431, 76.838
Kazakhstan_Medieval	NEO901	Sjauke settlement	782		D4e4	52.43, 76.839

[Kapisa]

TABLE S1
Sample ID: Site: Country: MT Haplogroup: y-chr Haplogroup: Age Average**: Data Source:
NEO281*:Kotias Klde: Georgia: H13c: J2b: 9,723 BP : This study
NEO283: Kotias Klde: Georgia: U4'9: - : 25,635 BP: This study
NEO310:Monjukli-Depe: Turkmenistan: H14: L1a: 6,550 BP: This study
NEO816*:Tepe Guran:Iran:R2: J2a: 8,700 BP : This study
NEO817:Tepe Guran:Iran: H2a2a1: - : 9,392 BP: This study
NEO819:Tepe Guran:Iran:U7a: - : 9,291 BP: This study

* Represent merged files due to identical genetic profiles. See "relatedness and notes" column, and see Table II for information on individual (non-merged) NEO-samples.
** Table VII

[RCO]

They are just refining and detailing the known genetic frontiers from the big four major ancestry clusters from the Mesolithic and the subsequent movements and dispersions. More samples and details will be found in other articles.

1288 Extended Data Fig 6. Spatiotemporal kriging of four major ancestry clusters over the last 12,000 years of human history.
1289 LVN = ancestry maximized in Anatolian farmer populations. WHG = ancestry maximized in western European hunter
1290 gatherers. EHG = ancestry maximized in eastern European hunter-gatherers. IRN = ancestry maximized in Iranian
1291 Neolithic individuals and Caucasus hunter-gatherers.